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Their homeworld was long since lost. Since circa 9,700 B.C, the remaining population of more than one million Fiilestii have resided in the Web of Worlds in a biome module customized for their physiology. It was spacious, airy, with kilometers-tall, spindle-like towers scattered throughout.
In 2269, Hikaru Sulu and Pavel Chekov explored the Fiilestii low-gravity module while wearing wingsuits to enable them to glide. When shifting gravitational conditions destabilized the module, the USS Enterprise helped evacuate citizens and tourists, but at least 3,600 people could not be saved. (TOS novel: The Face of the Unknown)